Cocos L., Sp. Pl. : 1188 (1753)

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Distribution

Map uses TDWG level 3 distributions (http://www.nhm.ac.uk/hosted_sites/tdwg/geogrphy.html)
Aldabrapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Andaman Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Angolapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Arubapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Assampresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Bahamaspresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Bangladeshpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Belizepresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Beninpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Bismarck Archipelagopresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Boliviapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Borneopresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Botswanapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Brazil Northpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Brazil Northeastpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Brazil Southeastpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Brazil West-Centralpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Burkinapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Burundipresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Cabindapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Cambodiapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Cameroonpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Cape Provincespresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Cape Verdepresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Caprivi Strippresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Caroline Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Cayman Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Central African Republicpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Chadpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
China South-Centralpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
China Southeastpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Christmas I.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Cocos (Keeling) Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Colombiapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Comorospresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Congopresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Cook Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Costa Ricapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Cubapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Dominican Republicpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Ecuadorpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
El Salvadorpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Equatorial Guineapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Ethiopiapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Fijipresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Floridapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Free Statepresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
French Guianapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Gabonpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Galápagospresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Gambia, Thepresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Ghanapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Gilbert Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Guatemalapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Guineapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Guinea-Bissaupresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Gulf of Guinea Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Guyanapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Hainanpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Haitipresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Hawaiipresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Honduraspresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Howland-Baker Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Indiapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Ivory Coastpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Jamaicapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Jawapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Kenyapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
KwaZulu-Natalpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Laccadive Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Laospresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Leeward Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Lesser Sunda Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Liberiapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Line Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Madagascarpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Malawipresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Malayapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Maldivespresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Malipresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Malukupresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Marcus I.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Marianaspresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Marquesaspresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Marshall Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Mauritaniapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Mauritiuspresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Mexican Pacific Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Mexico Centralpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Mexico Gulfpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Mexico Northeastpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Mexico Northwestpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Mexico Southeastpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Mexico Southwestpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Mozambiquepresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Mozambique Channel Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Myanmarpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Namibiapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Naurupresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Netherlands Antillespresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
New Caledoniapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
New Guineapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Nicaraguapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Nicobar Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Nigeriapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Niuepresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Northern Provincespresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Northern Territorypresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Panamápresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Perupresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Philippinespresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Phoenix Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Pitcairn Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Puerto Ricopresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Queenslandpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Rodriguespresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Rwandapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Réunionpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Samoapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Santa Cruz Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Senegalpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Seychellespresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Sierra Leonepresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Solomon Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Somaliapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
South China Seapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Southwest Caribbeanpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Sri Lankapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Sudanpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Sulawesipresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Sumaterapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Surinamepresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Swazilandpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Tanzaniapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Thailandpresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Togopresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Tokelau-Manihikipresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Tongapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Trinidad-Tobagopresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Tuamotupresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Tubuai Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Turks-Caicos Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Tuvalupresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Ugandapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Vanuatupresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Venezuelapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Venezuelan Antillespresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Vietnampresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Wake I.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Wallis-Futuna Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Western Australiapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Windward Is.present (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Zairepresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Zambiapresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
Zimbabwepresent (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)C
A single species widely cultivated throughout the tropics and warmer subtropics. Origin uncertain but said to be western Pacific (Harries 1978, Gruezo and Harries 1984, Buckley and Harries 1984) (but see below). (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)B

Belize, Benin, Bismarck Archipelago, Borneo, Caroline Is., Central American Pacific Is., Chile North, Costa Rica, Fiji, Gabon, Gilbert Is., Hawaii, India, Jawa, Leeward Is., Line Is., Madagascar, Malaya, Maldives, Maluku, Marcus I., Marianas, Marquesas, Marshall Is., Mauritius, Nauru, New Guinea, Nicaragua, Niue, Ogasawara-shoto, Philippines, Phoenix Is., Puerto Rico, Queensland, Réunion, Samoa, Seychelles, Society Is., Solomon Is., Sri Lanka, Thailand, Togo, Tokelau-Manihiki, Trinidad-Tobago, Tuamotu, Tubuai Is., Tuvalu, Vanuatu, Wallis-Futuna Is. (Govaerts, R. & Dransfield, J., World Checklist of Palms)A

Discussion

  • Cocos differs from other genera in Attaleinae in having large pistillate flowers with rounded sepals and petals, in the large fruit with thick fibrous mesocarp, and in the endosperm. Gruezo and Harries (1984) and Buckley and Harries (1984) record the presence of “wild-type” coconuts in apparently natural coastal forest in the Philippines and Australia, and argue that these areas are exactly where Cocos might be predicted to be native. Gunn (2004), however, in her phylogenetic analysis of Cocoseae, suggests a South American origin of the lineage that eventually gave rise to the coconut (see Chapter 7). (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)B

Diagnosis

  • The often slanting stems and graceful crowns of the coconut are largely responsible for palms being considered the hallmark of the tropics. Furthermore, the coconut, one of the ten most important crop trees, is the mainstay of many people. (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)B

Biology And Ecology

  • Cocos nucifera is often regarded as a strand plant but it will flower and fruit in humid equatorial regions at altitudes up to 900 m above sea level. Its natural habitat may well have been strand vegetation. (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)B

Common Name

  • Coconut (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)B

Etymology

  • Latinization of the Portuguese word, coco, originally used for a bugbear or ape, in reference to the face-like appearance of the partially dehusked endocarp. (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)B

Uses

  • One of the most important tropical crops with a multiplicity of uses both local and commercial. (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)B

Description

  • Moderate, solitary, unarmed, pleonanthic, monoecious palm, sometimes flowering while still without an emergent trunk. Stem erect, often curved or slanting, becoming bare and conspicuously ringed with leaf scars. Leaves numerous, pinnate, neatly abscising; sheath fibrous, forming a woven supportive network with a conspicuous, tongue-like extension opposite the petiole, eventually disintegrating and becoming open; petiole short to long, adaxially channelled, abaxially rounded, bearing caducous tomentum abaxially; rachis elongate, curved or straight, adaxially angled near the tip, abaxially rounded, with caducous tomentum abaxially; leaflets very numerous, single-fold, regularly arranged in one plane, usually rather stiff, linear, acuminate, usually bifid with slightly asymmetrical tips, adaxially glabrous, abaxially with abundant, dot-like scales and very small ramenta along the midrib, midrib prominent adaxially, transverse veinlets evident. Inflorescences solitary, interfoliar, axillary, branched to 1 order, protandrous; peduncle ± elliptic in cross-section, robust, elongate, bearing scattered scales; prophyll tubular, 2-keeled laterally, opening apically, becoming fibrous, tomentose, persistent, ± obscured by the leaf sheaths; peduncular bract inserted near the prophyll, very large, tubular, entirely enclosing the inflorescence until shortly before anthesis, splitting abaxially, becoming boat-shaped, beaked, thick, woody, adaxially smooth, abaxially with longitudinal, shallow grooves and caducous tomentum; rachis ±equalling the peduncle, bearing spirally arranged rachillae, each subtended by an inconspicuous triangular bract and with a swollen base; rachillae robust, ± pendulous at first, later spreading with a basal bare portion and none or a few basal triads and pairs or solitary staminate flowers distally; rachilla bracts and floral bracteoles inconspicuous. Staminate flowers ± asymmetrical, narrowly ovoid, moderate, sessile; sepals 3, distinct, rather unequal, imbricate, triangular, ± keeled; petals much longer than the sepals, thick, rather leathery, distinct, valvate, irregularly boat-shaped, acute; stamens 6, filaments rather short, distinct, awl-shaped, fleshy, ± erect, anthers deeply sagittate basally, shallowly so at the apex, elongate, medifixed, ± versatile, latrorse; pistillode with 3, slender, pointed lobes. Pollen ellipsoidal, frequently elongate and/or pyriform, usually with either slight or obvious asymmetry; aperture a distal sulcus; ectexine tectate, finely perforate, perforate and micro-channelled, or perforate-rugulate, aperture margin slightly finer; infratectum columellate; longest axis 62–70 µm [1/1]. Pistillate flowers very large, globose in bud, becoming very broadly ovoid at anthesis; sepals 3, distinct, imbricate, ± rounded; petals similar to and somewhat longer than the sepals, lacking valvate apices, very leathery; staminodal ring low, membranous, not lobed; gynoecium trilocular at the very base, triovulate, broadly ovoid, obscurely 3-angled, extremely fibrous distally, stigmas 3, very short, borne in a slight depression, ovule anatropous, very small, laterally attached. Fruit very large (except in unusual forms), ellipsoidal to broadly ovoid, indistinctly 3-angled, dull green, brown, brilliant-orange, yellow, to ivory-coloured when ripe, perianth enlarging in fruit, stigmatic remains apical; epicarp smooth, mesocarp very thick and fibrous, dry, endocarp thick and woody, ± spherical to narrow ovoid, indistinctly 3-angled, with 3 longitudinal ridges, and 3, large, slightly sunken, basal pores, each with an operculum. Seed almost always 1 only, very large, with a narrow layer of homogeneous endosperm, and a large central cavity partially filled with fluid; embryo basal, opposite one of the endocarp pores. Germination adjacent-ligular; eophyll entire, broadly lanceolate. Cytology: 2n = 32. (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)B

Anatomy

  • Leaf, stems, root (Tomlinson 1961), phloem (Parthasarathy 1974, 1980), wood (Chen 1995), root (Seubert 1998a, 1998b), megasporogenesis (Reddy and Kulkarni 1989), fruit (Roth 1977, Reddy and Kulkarni 1985). (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)B

Fossil record

  • Fruit or, more often, endocarps are the most frequently recorded fossils. The endocarps are distinguished by usually 3, sometimes more, well-defined pores. However, encocarps as large as those of Cocos nucifera are unknown. This suggests that the fossils may represent other genera within subtribe Attaleinae. Berry (1926b) first described Cocos zeylandica, a small endocarp (ca. 3.5 cm long), from Pliocene brown coal Cocos-bearing Beds in New Zealand (Mangonui, North Island); it was one of a number of specimens recovered. The endocarps are usually well-preserved and commonly washed up on the beach. Couper (1952), in his record of the pollen and spores of the Cocos-bearing Beds, notes that the rich flora of these beds had been recognised by geologists as early as 1872. From two pollen- and spore-rich samples, Couper (1952) records 4% and 8% abundance for Rhopalostylis/Cocos-type pollen (the two genera have closely similar pollen). Couper (1953) claims there can be little doubt that the commonly encountered palm pollen in the Cocos-bearing Beds is associated with Cocos zeylandica. However, although pollen of the two genera is closely similar in exine and size range (60–70 µm), whereas the pollen of other cocosoid palms with similar exine characteristics is smaller (<58 µm), it seems more reasonable to associate the fossil pollen with Rhopalostylis, a New Zealand palm. Further specimens of carbonised Cocos zeylandica were recovered from Miocene turbidites (possibly the result of a tsunami) at a different site in North Island by Ballance et al. (1981). From South Island, New Zealand (Otago Peninsula and Canterbury), Campbell et al. (2000) describe fossil endocarps (5–14.5 cm long) that have distinctive cocosoid pores, with the records from South Island ranging from mid-Eocene (45 million years ago) to Lower Oligocene (ca. 35 million years ago). Other records of fruits and seeds include silicified fruit (10 cm long x up to 9.5 cm wide) from the latest Pliocene of Australia (Queensland), which Rigby (1995) considers most resemble Cocos nucifera. Palmocarpon cetera, recorded from the Middle Oligocene of Puerto Rico, is compared with Cocos and Astrocaryum, although there is insufficient detail to make a satisfactory comparison (Hollick 1928). From the upper Cenomanien of France, Sainte Menehould area, Fliche (1894) describes an endocarp that bears a strong resemblance to cocosoid endocarps (6 cm in length, a thick testa 8 mm), to which he gives the name Cocoopsis. He also describes Cocoopsis ovata and C. zeilleri endocarps from the Lower Cenomanian of France (Fliche 1896); these have a diameter ranging from 41–60 mm, with a thick testa (8–12 mm). From India, Eocene (Rajasthan State), Cocos sahnii was described by Kaul (1951); large Cocos nucifera-like petrified fruit, 13 x 10 x 6 cm, with a thin epicarp, wide and fibrous mesocarp, and well-developed hardened endocarp are described from the Deccan Intertrappean deposits of Amarkantak, Madhya Pradesh (Tripathi et al. 1999) (although the age span of these volcanic deposits is controversial, see Chapter 5). A much smaller oval fruit (5 x 3 cm) with a smooth epicarp, fibrous mesocarp and stony endocarp, Cocos intertrappeansis, is recorded from the Tertiary of Mohgaonkalan (Madhya Pradesh) (Patil and Uphadhye 1984). An almost-complete Cocos-like palm stem, Palmoxylon sundaram, has been described from the Deccan Intertrappean beds of India by Sahni (1946). Although stem wood is difficult to identify to generic level, this is an interesting fossil and probably worth further study. Mahabalé (1978) discusses the fossil history and origins of the coconut. Harries (1978) comments, “... if the smaller-fruited coconuts identified in the New Hebrides Islands are taken into consideration, together with the (small) coconut fossil(s) in New Zealand, it may be suggested that the centre of origin was in the region of the submerged continental fragment of the Lord Howe Rise–Norfolk Ridge complex ... isolated from Australia about 80 my ago and apparently submerged below sea level about 15 my ago ...” However, it must be emphasised that determining the generic affinity of these fossils on the basis of their endocarps alone is problematic, as most of the genera of the Attaleinae are distinguished on morphological features unlikely to be preserved in the fossil record. (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)B

Relationships

  • Cocos is moderately supported as sister to Parajubaea (Baker et al. in review). (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)B

Taxonomic accounts

  • Glassman (1987) and Harries (1978, 1992). (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)B

Use Record

  • Cocos L.: Crop in garden of household. (Salick, J., Ecological basis of Amuesha agriculture, Peruvian upper Amazon. 1989)
    Use CategoryUse Sub CategoryPlant PartHuman GroupEthnic GroupCountry
    EnvironmentalAgroforestryEntire plantIndigenousYaneshaPeru

Bibliography

A. Govaerts, R. & Dransfield, J., World Checklist of Palms
B. J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008
C. J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008